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Dryopteris sandwicensis

(Hook. & Arn.) C.Chr. (1905)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Dryopteridaceae Genus: Dryopteris

dryopteris

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Bibliography

Name Published In: Index Filic.: 290 (1905)

Other References

Palmer 2003:147 (K, O, Mo, L, M, H); Degener 35262 BISH (L†, 1963)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Dryopteris sandwicensis Hanalei District; end of Kumuwela Ridge, gulches of Kokee stream, Kaou area Metrosideros-Acacia mesic forest with Alphitonia ponderosa, Santalum pyrularium, Leptocephylla tameamea, Coprosma waimeae, Dodonaea viscosa, Wikstroemia furcata, Melicope ovata, M. barbigera, Cyanea leptostegia Preserved_Specimen terrestrial rhizome erect, 5 cm long, scales shiny brown-red, associated with D. glabra, D. fusco-atra, D. unidentata, occasional Wood, K.R. 15197 Kauai BISH 2012-08-23
2 Dryopteris sandwicensis Halawa Mossy base of cliff near Moaula Fall. Preserved_Specimen St.John, H. 12735 Molokai BISH 1932-12-27
3 Dryopteris sandwicensis Waimea Canyon State Park. Puu Ka Pele Forest Reserve on western side of Waimea Canyon, just east of hwy 550 Steep slopes and open grassy areas, with remnants of dry mixed mesophytic forest of Metrosideros, Myrsine, Acacia, Cryptocarya and Zanthoxylum. Preserved_Specimen Common on open, steep, grassy slope with scattered Metrosideros and Nestegis. Flynn, T.W. 2738 Kauai BISH 1988-02-02
4 Dryopteris sandwicensis W Maui, Lahaina District, Lihau Lowland Koa Mesic Forest at 3370 ft under a canopy of Acacia koa, Psychotria mariniana, Myrsine, Coprosma, Dodonaea and Metrosideros with a variety of native ferns, sedges and grasses Preserved_Specimen Welton, P. 1323 Maui BISH 1991-08-28
5 Dryopteris sandwicensis Waialae drainage, moderate to steep banks above stream. Acacia-Metrosideros mesic forest, open to closed canopy. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Syzygium snadwicensis, Perrottetia sandwicensis, Psychotria mariniana, Bidens cosmoides, Pouteria, Xylosma hawaiiense, Alyxia, Chamaesyce atrococ Preserved_Specimen terrestrial rhizome erect or sub-erect 10 cm. x 10 cm., scales dark brown at base of stipe. Wood, K.R. 13830 Kauai BISH 2009-07-17
6 Dryopteris sandwicensis Waianae District: Waianae Mts., near beginning of first gulch below Puu Hapapa, just off Honouliuli Contour Trail Wet, shaded; associated with Cyrtandra, Charpentiera, Urera, ferns. Preserved_Specimen Herbst, D.R. 420 Oahu BISH 1967-04-15
7 Dryopteris sandwicensis Waianae Mountains, Puu Kaala above Waialua on the DuPonte ridge trail Forest on steep ridge. Preserved_Specimen Stone, B.C. 3169 Oahu BISH 1959-12-20
8 Dryopteris sandwicensis Forests. Preserved_Specimen Nelson, D. 22 Hawaii BISH 1779-01-01
9 Dryopteris sandwicensis Waianae Range; Mokuleia. West branch, East Makaleha Valley, in small side gulch Occasional in dryland forest dominated by Diospyros ferrea var. sandwicensis. Preserved_Specimen Hatheway, W.H. 390 Oahu BISH 1950-09-30
10 Dryopteris sandwicensis Waikolu Valley Wet forest. Steep valley wall below Puu Kaeo. Preserved_Specimen St.John, H. 23452 Molokai BISH 1948-12-22
11 Dryopteris sandwicensis Koolau Mts., Punaluu. Preserved_Specimen Rock, J.F.C. 233 Oahu BISH 1908-11-14
12 Dryopteris sandwicensis Waiakeakua Preserved_Specimen Munro, G.C. 56 Lanai BISH 1927-06-07
13 Dryopteris sandwicensis W Maui, Lahaina District, Lihau Lowland Ohia Mixed Mesic Shrubland at 3400 ft in a gulch below Lihau with Metrosideros, Antidesma, Psychotria, Nestegis, Ilex, Cheirodendron, Myrsine, Xylosma, Pelea, Wikstroemia, Coprosma, Dodonaea, Styphelia, Sadleria and native ferns and sedges Preserved_Specimen Welton, P. 1495 Maui BISH 1992-01-10
14 Dryopteris sandwicensis East Maui, Olinda, on the side of a small gulch just above Waihou Spring Forest Reserve Preserved_Specimen Fern 2-2.5 ft tall; stipe sparsely clothed with 5-7 mm x 1-2 mm curly scales at base, becoming glabrous before reaching frond; stipe thrice sulcate; frond tripinnate, pinnules long and narrow, not chartaceous, veins in pinnules pinnate, ultimately once forking, distinct; sori naked; sprangia in round clusters. Hobdy, R.W. 1026 Maui BISH 1981-03-03
15 Dryopteris sandwicensis Monini Gulch next to an area recently burned. Preserved_Specimen Corn, C. Oahu BISH 1984-05-24
16 Dryopteris sandwicensis Lihue District: Hoary Head Range. North side of range between Haupu and Kamaulele Below low saddle along ridge in area dominated by Cordyline with Psidium and Metrosideros. Wet area w/number of ferns including Christella, Microlepia, Diplazium, Blechnum and Sphenomeris. Preserved_Specimen Flynn, T.W. 2945 Kauai BISH 1988-04-21
17 Dryopteris sandwicensis Kalaupapa; Waikolu, S, stream head of valley Preserved_Specimen medium terrestrial fern Wysong, M.L. 492 Molokai BISH 2004-11-23
18 Dryopteris sandwicensis Koele, mountains near Preserved_Specimen Forbes, C.N. 81 Lanai BISH 1913-06-01
19 Dryopteris sandwicensis W Maui District, Pohakea Gulch, Hanaula Preserved_Specimen Hobdy, R.W. 531 Maui BISH 1979-05-10
20 Dryopteris sandwicensis Lihue District: Hoary Head Range. North side of range between Haupu and Kamaulele Below low saddle along ridge in area dominated by Cordyline with Psidium and Metrosideros. Wet area w/number of ferns including Christella, Microlepia, Diplazium, Blechnum and Sphenomeris. Preserved_Specimen Flynn, T.W. 2945 Kauai BISH 1988-04-21
21 Dryopteris sandwicensis W Maui, Hanaula Springs Trail Preserved_Specimen Large fern; stipes 2.5-3 ft long, glabrous except for sparse scales near base; frond to 3.5 ft long, tripinnatifid; sori naked. Hobdy, R.W. 1066 Maui BISH 1981-05-06
22 Dryopteris sandwicensis West Maui; Olowalu, near head of valley, lateral ridge near center Preserved_Specimen Stipe 98 cm. long, frond? 116 cm. long, rhizome creeping. Forbes, C.N. 2422 Maui BISH 1920-05-19
23 Dryopteris sandwicensis Kaala Mountains Preserved_Specimen U.S. Exploring Expedition 28 Oahu BISH 1838-01-01
24 Dryopteris sandwicensis Waianae Mountains, Honouluili, Pohakea Pass Dry slopes. Preserved_Specimen Fosberg, F.R. 9508 Oahu BISH 1933-05-12
25 Dryopteris sandwicensis W Maui, Olowalu, near head of valley, lateral ridge near center Preserved_Specimen Stipe 98 cm. long, frond? 116 cm. long, rhizome creeping. Forbes, C.N. 2422 Maui BISH 1920-05-19
26 Dryopteris sandwicensis Kalanilehua, Olaa. Preserved_Specimen Tip of frond. Giffard, W.M. Hawaii BISH 1927-07-16
27 Dryopteris sandwicensis Kalanilehua, Olaa. Preserved_Specimen Basal pinna. Giffard, W.M. Hawaii BISH 1927-07-16
28 Dryopteris sandwicensis Waianae Mountains, Dupont Trail to Puu Kaala Preserved_Specimen Krajina, V.J. 620225335 Oahu BISH 1962-02-25
29 Dryopteris sandwicensis Olowalu Valley, central ridge at head, near backbone peak Preserved_Specimen Forbes, C.N. 2357 Maui BISH 1920-05-12
30 Dryopteris sandwicensis East Maui, Koolau Forest Reserve (now Hanawi Natural Area Reserve) Preserved_Specimen Gagné, B.H. 353 Maui BISH 1973-07-22
31 Dryopteris sandwicensis East Maui, Waiopae Gulch, growing at Kahikinui Preserved_Specimen Hobdy, R.W. 3039 Maui BISH 1989-05-04
32 Dryopteris sandwicensis unknown Preserved_Specimen Mann, H. 592 BISH 1864-01-01
33 Dryopteris sandwicensis unknown Preserved_Specimen Whitney, J.M. BISH 1870-01-01
34 Dryopteris sandwicensis Keawapilau Gulch Preserved_Specimen Welton, P. 638 Oahu BISH 1991-01-20
35 Dryopteris sandwicensis Ridge east of Kuliououiki Preserved_Specimen Forbes, C.N. 2013 Oahu BISH 1914-11-17
36 Dryopteris sandwicensis W Maui, Hanaula Springs Trail Preserved_Specimen Large fern; stipes 2.5-3 ft long, glabrous except for sparse scales near base; frond to 3.5 ft long, tripinnatifid; sori naked. Hobdy, R.W. 1066 Maui BISH 1981-05-06
37 Dryopteris sandwicensis East Maui, Waiopae Gulch, growing at Kahikinui Preserved_Specimen Hobdy, R.W. 3039 Maui BISH 1989-05-04
38 Dryopteris sandwicensis Maui Preserved_Specimen Mann, H. 592 Maui BISH 1864-01-01
39 Dryopteris sandwicensis In the high gulches of Puu Kawiwi, west of the Waianae Kai road Preserved_Specimen Palmer, D.D. 765 Oahu BISH 1991-12-15
40 Dryopteris sandwicensis unknown Preserved_Specimen BISH
41 Dryopteris sandwicensis unknown Preserved_Specimen (Not on sheet) 880 BISH
42 Dryopteris sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
43 Dryopteris sandwicensis Waianae Mountains, Maunakapu-Palikea Preserved_Specimen Krajina, V.J. 611029030 Oahu BISH 1961-10-29
44 Dryopteris sandwicensis Haleakala, Keanae Gap, Halehoku Preserved_Specimen Forbes, C.N. 1077 Maui BISH 1919-08-07
45 Dryopteris sandwicensis Kahanui Preserved_Specimen Krajina, V.J. 620612069 Molokai BISH 1962-06-12
46 Dryopteris sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 532 Lanai BISH 1916-03-25
47 Dryopteris sandwicensis Between the Hanapepe and Wahiawa Rivers Preserved_Specimen Heller, A.A. 2587 Kauai BISH 1895-07-18
48 Dryopteris sandwicensis Waihou Spring, Olinda Preserved_Specimen Hobdy, R.W. 356 Maui BISH 1978-10-23
49 Dryopteris sandwicensis Maui Preserved_Specimen Mann, H. 592 Maui BISH 1864-01-01
50 Dryopteris sandwicensis Haupu Range. Left hand side of Kipu Kai Gap Preserved_Specimen Forbes, C.N. 723 Kauai BISH 1916-11-01
51 Dryopteris sandwicensis Koele, mountains near Preserved_Specimen Forbes, C.N. 81 Lanai BISH 1913-06-01
52 Dryopteris sandwicensis Maui Preserved_Specimen Mann, H. 591 Maui BISH 1864-01-01
53 Dryopteris sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 477 Lanai BISH 1915-12-08
54 Dryopteris sandwicensis Mokuleia Trail Preserved_Specimen Welton, P. 746 Oahu BISH 1991-07-02
55 Dryopteris sandwicensis West Maui, summit forests near Puu Kukui Trail. Preserved_Specimen Hobdy, R.W. 3233 Maui BISH 1991-07-31
56 Dryopteris sandwicensis Return from Mauna Kea Preserved_Specimen U.S. Exploring Expedition 26 Hawaii BISH 1840-01-01
57 Dryopteris sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 167 Lanai BISH 1913-10-18
58 Dryopteris sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 167 Lanai BISH 1913-10-18
59 Dryopteris sandwicensis Makua Preserved_Specimen Russ, G.W. Oahu BISH 1930-05-01
60 Dryopteris sandwicensis Wailupe Valley, right fork Preserved_Specimen Garber, D.W. 160 Oahu BISH 1920-01-12
61 Dryopteris sandwicensis Wailupe Valley, right fork Preserved_Specimen Garber, D.W. 160 Oahu BISH 1920-01-12
62 Dryopteris sandwicensis unknown Preserved_Specimen Flynn, T.W. 2757 Kauai BISH 1988-02-17
63 Dryopteris sandwicensis Between the Hanapepe and Wahiawa Rivers Preserved_Specimen Heller, A.A. 2587 Kauai BISH 1895-07-18
64 Dryopteris sandwicensis Waiakeakua Preserved_Specimen Munro, G.C. 90 Lanai BISH 1927-06-14
65 Dryopteris sandwicensis Haupu Range. Left hand side of Kipu Kai Gap Preserved_Specimen Forbes, C.N. 723 Kauai BISH 1916-11-01
66 Dryopteris sandwicensis On dry lava beds back of Huehue on the way to Puuwaawaa Preserved_Specimen Rock, J.F.C. 3625 Hawaii BISH 1909-06-04
67 Dryopteris sandwicensis Kaohai, Awehi Gulch, head of, in lower forest Preserved_Specimen St.John, H. 18881 Lanai BISH 1938-04-15
68 Dryopteris sandwicensis Haleakala, Keanae Gap, Halehoku Preserved_Specimen Forbes, C.N. 1077 Maui BISH 1919-08-07
69 Dryopteris sandwicensis Oahu Preserved_Specimen Mann, H. 185 Oahu BISH 1864-01-01
70 Dryopteris sandwicensis Forest of Waimea Preserved_Specimen Lyons, A.B. Hawaii BISH 1860-01-01
71 Dryopteris sandwicensis Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-22
72 Dryopteris sandwicensis Forest of Waimea Preserved_Specimen Lyons, A.B. Hawaii BISH 1860-01-01
73 Dryopteris sandwicensis Hanapepe Preserved_Specimen Wiebke, H. Kauai BISH 1926-06-19
74 Dryopteris sandwicensis Oahu Preserved_Specimen Beechey, Capt. F.W. Oahu BISH
75 Dryopteris sandwicensis unknown Preserved_Specimen Flynn, T.W. 2111 BISH 1987-03-20
76 Dryopteris sandwicensis unknown Preserved_Specimen Baldwin, D.D. BISH 1879-08-14
77 Dryopteris sandwicensis unknown Preserved_Specimen Lack, F. BISH 1880-01-01
78 Dryopteris sandwicensis East Maui, found above Kahipa's Place Preserved_Specimen (Not on sheet) Maui BISH
79 Dryopteris sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
80 Dryopteris sandwicensis Maui Preserved_Specimen Mann, H. 592 Maui BISH 1864-01-01
81 Dryopteris sandwicensis Waianae Mts.; Maunakapu-Palikea 21.4N 158.1W Preserved_Specimen Krajina, V.J. 611029030 Oahu BISH 1961-10-29
82 Dryopteris sandwicensis Forest above Saw Mill Preserved_Specimen U.S. Exploring Expedition 29 Hawaii BISH 1840-01-01
83 Dryopteris sandwicensis unknown Preserved_Specimen Rock, J.F.C. BISH
84 Dryopteris sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 411 Lanai BISH 1915-12-08
85 Dryopteris sandwicensis Pahole Drainage #2 - releve 18 Preserved_Specimen Welton, P. 573 Oahu BISH 1990-07-09
86 Dryopteris sandwicensis All the Islands Preserved_Specimen BISH
87 Dryopteris sandwicensis unknown Preserved_Specimen BISH
88 Dryopteris sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
89 Dryopteris sandwicensis W Maui, Lahaina District, Lihau Ohia Mixed Mesic Forest at 3700 ft with Metrosideros, Myrsine, Dodonaea, Coprosma, Lythrum, Lysimachia, Vaccinium, Polypodium, Doodia kunthiana, and Elaphoglossum Preserved_Specimen Welton, P. 992 Maui BISH 1991-07-09
90 Dryopteris sandwicensis N (mauka) of Lanai City, N (mauka) In patch of decadent forest. Preserved_Specimen Degener, O. 22017 Lanai BISH 1952-03-02
91 Dryopteris sandwicensis West Maui, Hanaula, upper section of Pohakea Gulch, west of parking area at base of springs trail Aspect: W. Scattered plants on slopes of gulch, mixed in with native understorey and Melinus. Mesic: Metrosideros, Cheirodendron, Myrlsine, Coprosma, Notelaea, Cyrtandra, Sadleria, Nephrolepis, Microlepia, Erigeron, Adiatum, Melinus Preserved_Specimen Pezzillo, Z. 1089 Maui BISH 2024-12-12
92 Dryopteris sandwicensis Lahaina District, Main (left) fork of Manawainui Gulch In shade of Eugenia and Metrosideros w/grasses and Alyxia. Preserved_Specimen Rootstock erect to oblique, rising 8-11 cm above ground, 8 cm thick. Nagata, K.M. 1914 Maui BISH 1979-06-06
93 Dryopteris sandwicensis Nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, W/ Alphitonia ponderosa, Claoxylon sandwicensis, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea viscosa, Kadua affinis, Melicope ovata, Pleomele aurea, Tetraplasandra kavaiensis, Psychotria green we Preserved_Specimen terrestrial rhizome sub-erect, exindusiate, scales dark brown stiff at base of stipe diminishing up into rachis, occasional Wood, K.R. 14522 Kauai BISH 2011-02-19
94 Dryopteris sandwicensis Waianae Mountains; Pohakea Pass, Halona, Lualualei Wooded slope. Preserved_Specimen St.John, H. 13134 Oahu BISH 1933-05-12
95 Dryopteris sandwicensis Hapapa, crest of approach ridge In grassy clearing beneath Grevillea. Preserved_Specimen Scattered clumps. Paquin, D. 3498 Oahu BISH 1987-03-28
96 Dryopteris sandwicensis Waimea District. Kohua Ridge and south facing slopes above Koaie. Metrosideros - Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bob Preserved_Specimen Terrestrial fern, rhizomes sub-erect, 7 cm long x 5 cm wide, ca. 7 fronds, occasional Wood, K.R. 17157 Kauai BISH 2016-10-20
97 Dryopteris sandwicensis Kohua Ridge and south facing slopes above Koaie. Metrosideris-Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bob Preserved_Specimen Terrestrial fern, rhizomes sub-erect, 7 cm long x 5 cm wide, ca. 7 fronds, occasional. Wood, K.R. 17157 Kauai BISH 2016-10-20
98 Dryopteris sandwicensis On Waianae Range. On summit of Mt. Kaala Scattered. Preserved_Specimen Herat, T. 528 Oahu BISH 1973-09-30
99 Dryopteris sandwicensis Waianae Ditch Trail Shady Bank. Preserved_Specimen Topping, D.L. 3611 Oahu BISH 1932-09-11
100 Dryopteris sandwicensis Nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, W/ Alphitonia ponderosa, Claoxylon sandwicensis, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea viscosa, Kadua affinis, Melicope ovata, Pleomele aurea, Tetraplasandra kavaiensis, Psychotria green we Preserved_Specimen terrestrial rhizome sub-erect, exindusiate, scales dark brown stiff at base of stipe diminishing up into rachis, occasional Wood, K.R. 14522 Kauai BISH 2011-02-19
101 Dryopteris sandwicensis East Maui, Auwahi, SE Puu Ouli, S. slope of Haleakala Rare on ledges in dry open sclerophyllous forest on a`a lava flow. Preserved_Specimen Fosberg, F.R. 41506 Maui BISH 1961-03-14
102 Dryopteris sandwicensis North Fork of Kaluaa gulch, Waianae mountains. Growing in talus of gulch bottom in mesic forest dominated by Schinus, Fraxinus, Planchonella, Ceodes, Microlepia, Plagiochila, Gynocthodes, and others. Preserved_Specimen Semi erect fern with rhizome making a short trunk. Scales twisted and linear, congested at base of stipe. Scattered cauducous hairs along stipe, rachis, and costae but otherwise quite smooth and glabrous. Stipe straw colored. Sori mostly marginal to submarginal. Many Elasmius snails on certain fronds. Thomas, M.K. 286 Oahu BISH 2022-03-16
103 Dryopteris sandwicensis Hapapa Terrestrial off crest of buttress. Preserved_Specimen Takeuchi, W.N. 3500 Oahu BISH 1987-03-28
104 Dryopteris sandwicensis Kamakou Preserve, E side of Kawela Ridge, due SW of Puu o Kaeha Low wet forest w/Metrosideros dominant on lower slopes and valley bottom, with Styphelia/Dodonaea shrubland on upper slopes and ridgetops. Common on slopes with light shade. Preserved_Specimen ground fern with stout, suberect rhizome. Lorence, D.H. 5617 Molokai BISH 1987-10-21
105 Dryopteris sandwicensis Hapapa Terrestrial off crest of buttress. Preserved_Specimen Takeuchi, W.N. 3500 Oahu BISH 1987-03-28
106 Dryopteris sandwicensis Kalulu, Kapano Gulch Wooded gulch. Preserved_Specimen St.John, H. 18757 Lanai BISH 1938-04-10
107 Dryopteris sandwicensis Kau, Manuka, Govt. road Fern forest, shaded, dry. Preserved_Specimen Russ, G.W. 22 Hawaii BISH 1927-07-11
108 Dryopteris sandwicensis Kohua Ridge and south facing slopes above Koaie. Metrosideris-Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bob Preserved_Specimen Terrestrial fern, rhizomes sub-erect, 7 cm long x 5 cm wide, ca. 7 fronds, occasional. Wood, K.R. 17157 Kauai BISH 2016-10-20
109 Dryopteris sandwicensis East Maui, Hana District, S slope of Haleakala, Auwahi Forest Rare on side of tiny ravine in open dry sclerophyll forest on a`a lava. Preserved_Specimen Fosberg, F.R. 48336 Maui BISH 1967-03-25
110 Dryopteris sandwicensis Nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, W/ Alphitonia ponderosa, Claoxylon sandwicensis, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea viscosa, Kadua affinis, Melicope ovata, Pleomele aurea, Tetraplasandra kavaiensis, Psychotria greenwe Preserved_Specimen terrestrial rhizome sub-erect, exindusiate, scales dark brown stiff at base of stipe diminishing up into rachis, occasional Wood, K.R. 14522 Kauai BISH 2011-02-19
111 Dryopteris sandwicensis Palawai, Lanaihale Tufts in moist forest. Preserved_Specimen St.John, H. 18865 Lanai BISH 1938-04-15
112 Dryopteris sandwicensis Makua Valley Shady gully Preserved_Specimen Topping, D.L. 3556 Oahu BISH 1932-06-26
113 Dryopteris sandwicensis Waihou Gulch in pasture below Olinda reservoir On a damp bank out of reach of the cattle. Preserved_Specimen 2 foot fern, stipe scaly only in lower portion, scales broad stramineous, sori naked. Hobdy, R.W. 765 Maui BISH 1980-02-03
114 Dryopteris sandwicensis Puu Kaua, east ridge, Honouliuli, Waianae Mts Dry forest. Preserved_Specimen Higgins, N.H. 63 Oahu BISH 1955-12-11
115 Dryopteris sandwicensis back of Huehue on road to Puuwaawaa dry lava fields Preserved_Specimen Rock, J.F.C. 3631 Hawaii BISH 1909-06-01
116 Dryopteris sandwicensis Waikolu Valley Wet forest. Steep valley wall below Puu Kaeo. Preserved_Specimen St.John, H. 23452 Molokai BISH 1948-12-22
117 Dryopteris sandwicensis Hapapa Terrestrial off crest of buttress. Preserved_Specimen Takeuchi, W.N. 3500 Oahu BISH 1987-03-28
118 Dryopteris sandwicensis North Fork of Kaluaa gulch, Waianae mountains. Growing in talus of gulch bottom in mesic forest dominated by Schinus, Fraxinus, Planchonella, Ceodes, Microlepia, Plagiochila, Gynocthodes, and others. Preserved_Specimen Semi erect fern with rhizome making a short trunk. Scales twisted and linear, congested at base of stipe. Scattered cauducous hairs along stipe, rachis, and costae but otherwise quite smooth and glabrous. Stipe straw colored. Sori mostly marginal to submarginal. Many Elasmius snails on certain fronds. Thomas, M.K. 286 Oahu BISH 2022-03-16
119 Dryopteris sandwicensis Palawai, Lanaihale Tufts in moist forest. Preserved_Specimen St.John, H. 18865 Lanai BISH 1938-04-15
120 Dryopteris sandwicensis Koolau Range, Kipapa-Waiahole Crest Metrosideros-Cheirodendron forest. Preserved_Specimen Grant, M.L. 7230 Oahu BISH 1934-08-07
121 Dryopteris sandwicensis Waikolu Valley Wet forest. Steep valley wall below Puu Kaeo. Preserved_Specimen St.John, H. 23452 Molokai BISH 1948-12-22
122 Dryopteris sandwicensis Kapano Gulch Partly shaded slope. Preserved_Specimen Degener, O. 35262 Lanai BISH 1963-08-31
123 Dryopteris sandwicensis W Maui, Lahaina District, Lihau Lowland Koa Mesic Forest at 3370 ft under a canopy of Acacia koa, Psychotria mariniana, Myrsine, Coprosma, Dodonaea and Metrosideros with a variety of native ferns, sedges and grasses Preserved_Specimen Welton, P. 1324 Maui BISH 1991-08-28
124 Dryopteris sandwicensis Southeast slope of Puu Hapapa Dense forest. Preserved_Specimen Degener, O. 12351 Oahu BISH 1939-06-04
125 Dryopteris sandwicensis Kau, Manuka, forest reserve Dry, hot, a`a flow, with Nephrolepis exaltata Schott. Exposed to sun. Preserved_Specimen Height of plant 2-1/2 ft. Russ, G.W. 19 Hawaii BISH 1927-07-01
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